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In biology, evolution is the change in the heritable traits of a population over successive generations, as determined by the shifting allele frequencies of genes. Evolution is ultimately the source of the vast diversity of life: all contemporary organisms are related to each other through common descent, products of cumulative evolutionary changes over billions of years. Over time, new species evolve from existing species through speciation, and other species become extinct, resulting in the ever-changing biological world reflected in the fossil record.

The basic mechanisms that produce evolutionary change are natural selection (which includes ecological and sexual selection) and genetic drift acting on the genetic variation created by mutation, genetic recombination and gene flow. Natural selection is the process by which individual organisms with favorable traits are more likely to survive and reproduce. If those traits are heritable, they pass them to their offspring, with the result that beneficial heritable traits become more common in the next generation. Given enough time, this passive process can result in varied adaptations to changing environmental conditions.

The modern understanding of evolution is based on the theory of natural selection, which was first set out in a joint 1858 paper by Charles Darwin and Alfred Russel Wallace and popularized in Darwin's 1859 book The Origin of Species. In the 1930s, scientists combined Darwinian natural selection with the theory of Mendelian heredity to form the modern evolutionary synthesis, also known as "Neo-Darwinism". The modern synthesis describes evolution as a change in the frequency of alleles within a population from one generation to the next. This theory has become the central organizing principle of modern biology, relating directly to topics such as the origin of antibiotic resistance in bacteria, eusociality in insects, and the staggering biodiversity of the living world.

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